Set in an alternate America where witches ended their persecution over 300 years ago by cutting a deal with the burgeoning government to fight for their country, the series follows three young women from basic training in combat magic into early deployment. In this world, the traditional roles of gender and power are flipped, with the more dominant women on the front lines fighting looming terrorist threats that are familiar to our world, but with supernatural tactics and weapons.

The unit travels to Salem Town for an annual witch celebration, but an unexpected disturbance throws the proceedings into chaos. Tally wonders if she made the right choice to embark on a military career when witch-hate and her own mother stand in her way. Smarting from Alder’s denial of her request to switch units, Abigail attempts to bury the hatchet with Raelle. Raelle grows closer to Scylla, while the unit impresses Anacostia in Windstrike training.

Fort Salem excitedly greets male visitors on the eve of Beltane, but Raelle and Scylla’s budding romance is interrupted by the arrival of a mysterious man from Scylla’s past. One of the male witches catches Tally’s eye, but she is unsure of how to proceed. Meanwhile, Alder attempts to convince the Hague, an international military council of witches, to follow her strategy against the Spree and is startled by the news of an escalating humanitarian crisis in the Tarim Basin.

The base reels from a traumatic event, while Raelle tries to come to terms with her role in the tragedy. Scylla attempts to evade Raelle’s increasingly pointed questions. Abigail works to impress her formidable rival, Libba. And Tally’s connection with Gerit grows during Beltane, a sacred holiday honoring fertility traditions.

The unit travels off base for the high society wedding of Abigail’s cousin. Raelle throws herself into her relationship with Scylla, who, unbeknownst to Raelle, faces an impossible choice. Abigail becomes disillusioned with her privilege while amongst the notable Bellweather line. And Tally gets hit with two life-altering revelations that change the very fabric of the unit.